Результаты поиска по 'heterogeneous population':
Найдено статей: 7
  1. Alpeeva L.E., Tsybulin V.G.
    The cosymmetric approach to the analysis of spatial structure of populations with amount of taxis
    Computer Research and Modeling, 2016, v. 8, no. 4, pp. 661-671

    We consider a mathematical model describing the competition for a heterogeneous resource of two populations on a one-dimensional area. Distribution of populations is governed by diffusion and directed migration, species growth obeys to the logistic law. We study the corresponding problem of nonlinear parabolic equations with variable coefficients (function of a resource, parameters of growth, diffusion and migration). Approach on the theory the cosymmetric dynamic systems of V. Yudovich is applied to the analysis of population patterns. Conditions on parameters for which the problem under investigation has nontrivial cosymmetry are analytically derived. Numerical experiment is used to find an emergence of continuous family of steady states when cosymmetry takes place. The numerical scheme is based on the finite-difference discretization in space using the balance method and integration on time by Runge-Kutta method. Impact of diffusive and migration parameters on scenarios of distribution of populations is studied. In the vicinity of the line, corresponding to cosymmetry, neutral curves for diffusive parameters are calculated. We present the mappings with areas of diffusive parameters which correspond to scenarios of coexistence and extinction of species. For a number of migration parameters and resource functions with one and two maxima the analysis of possible scenarios is carried out. Particularly, we found the areas of parameters for which the survival of each specie is determined by initial conditions. It should be noted that dynamics may be nontrivial: after starting decrease in densities of both species the growth of only one population takes place whenever another specie decreases. The analysis has shown that areas of the diffusive parameters corresponding to various scenarios of population patterns are grouped near the cosymmetry lines. The derived mappings allow to explain, in particular, effect of a survival of population due to increasing of diffusive mobility in case of starvation.

    Views (last year): 2. Citations: 1 (RSCI).
  2. Ha D.T., Tsybulin V.G.
    Diffusion–reaction–advection equations for the predator–prey system in a heterogeneous environment
    Computer Research and Modeling, 2021, v. 13, no. 6, pp. 1161-1176

    We analyze variants of considering the inhomogeneity of the environment in computer modeling of the dynamics of a predator and prey based on a system of reaction-diffusion–advection equations. The local interaction of species (reaction terms) is described by the logistic law for the prey and the Beddington –DeAngelis functional response, special cases of which are the Holling type II functional response and the Arditi – Ginzburg model. We consider a one-dimensional problem in space for a heterogeneous resource (carrying capacity) and three types of taxis (the prey to resource and from the predator, the predator to the prey). An analytical approach is used to study the stability of stationary solutions in the case of local interaction (diffusionless approach). We employ the method of lines to study diffusion and advective processes. A comparison of the critical values of the mortality parameter of predators is given. Analysis showed that at constant coefficients in the Beddington –DeAngelis model, critical values are variable along the spatial coordinate, while we do not observe this effect for the Arditi –Ginzburg model. We propose a modification of the reaction terms, which makes it possible to take into account the heterogeneity of the resource. Numerical results on the dynamics of species for large and small migration coefficients are presented, demonstrating a decrease in the influence of the species of local members on the emerging spatio-temporal distributions of populations. Bifurcation transitions are analyzed when changing the parameters of diffusion–advection and reaction terms.

  3. Gerasimov A.N., Shpitonkov M.I.
    Mathematical model of the parasite – host system with distributed immunity retention time
    Computer Research and Modeling, 2024, v. 16, no. 3, pp. 695-711

    The COVID-19 pandemic has caused increased interest in mathematical models of the epidemic process, since only statistical analysis of morbidity does not allow medium-term forecasting in a rapidly changing situation.

    Among the specific features of COVID-19 that need to be taken into account in mathematical models are the heterogeneity of the pathogen, repeated changes in the dominant variant of SARS-CoV-2, and the relative short duration of post-infectious immunity.

    In this regard, solutions to a system of differential equations for a SIR class model with a heterogeneous duration of post-infectious immunity were analytically studied, and numerical calculations were carried out for the dynamics of the system with an average duration of post-infectious immunity of the order of a year.

    For a SIR class model with a heterogeneous duration of post-infectious immunity, it was proven that any solution can be continued indefinitely in time in a positive direction without leaving the domain of definition of the system.

    For the contact number $R_0 \leqslant 1$, all solutions tend to a single trivial stationary solution with a zero share of infected people, and for $R_0 > 1$, in addition to the trivial solution, there is also a non-trivial stationary solution with non-zero shares of infected and susceptible people. The existence and uniqueness of a non-trivial stationary solution for $R_0 > 1$ was proven, and it was also proven that it is a global attractor.

    Also, for several variants of heterogeneity, the eigenvalues of the rate of exponential convergence of small deviations from a nontrivial stationary solution were calculated.

    It was found that for contact number values corresponding to COVID-19, the phase trajectory has the form of a twisting spiral with a period length of the order of a year.

    This corresponds to the real dynamics of the incidence of COVID-19, in which, after several months of increasing incidence, a period of falling begins. At the same time, a second wave of incidence of a smaller amplitude, as predicted by the model, was not observed, since during 2020–2023, approximately every six months, a new variant of SARS-CoV-2 appeared, which was more infectious than the previous one, as a result of which the new variant replaced the previous one and became dominant.

  4. Ainbinder R.M., Rassadin A.E.
    On population migration in an ecological niche with a spatially heterogeneous local capacity
    Computer Research and Modeling, 2025, v. 17, no. 3, pp. 483-500

    The article describes the migration process of a certain population, taking into account the spatial heterogeneity of the local capacity of the ecological niche. It is assumed that this spatial heterogeneity is caused by various natural or artificial factors. The mathematical model of the migration process under consideration is a Cauchy problem on a straight line for some quasi-linear partial differential equation of the first order, which is satisfied by the linear population density under consideration. In this paper, a general solution to this Cauchy problem is found for an arbitrary dependence of the local capacity of an ecological niche on the spatial coordinate. This general solution was applied to describe the migration of the population in question in two different cases: in the case of a dependence of the local capacity of the ecological niche on the spatial coordinate in the form of a smooth step and in the case of a hill-like dependence of the local capacity of the ecological niche on the spatial coordinate. In both cases, the solution to the Cauchy problem is expressed in terms of higher transcendental functions. By applying special relations to the model parameters, these higher transcendental functions are reduced to elementary functions, which makes it possible to obtain exact model solutions explicitly expressed in terms of elementary functions. With the help of these precise solutions, an extensive program of computational experiments has been implemented, showing how the initial population density of the Gaussian form is dispersed by the considered two types of spatial heterogeneity of the local capacity of the ecological niche. These computational experiments have shown that when passing through both step-like and hill-like spatial inhomogeneities of the local capacity of an ecological niche with a narrow Gaussian width of its initial density compared to the characteristic spatial scale of these inhomogeneities, the system forgets its initial state. In particular, if we interpret the system under study as a population living in an extended calm rectilinear river along its bed, then it can be argued that under this initial condition, after the current of this river carries the population under consideration through the area of spatial heterogeneity of the local capacity of the ecological niche, the population density becomes a quasi-rectangular function.

  5. Pirutin S.K., Shank M.A., Jia S., Konuhov I.V., Todorenko D.A., Chervitsov R.N., Fursova P.V., Kabashnikova L.F., Plusnina T.Yu., Khruschev S.S., Riznichenko G.Yu., Rubin A.B.
    Comprehensive analysis of copper ions effect on the primary processes of photosynthesis in Scenedesmus quadricauda based on chlorophyll a fluorescence measurements in suspension and on single cells
    Computer Research and Modeling, 2025, v. 17, no. 2, pp. 293-322

    The effect of copper ions on the primary processes of photosynthesis in freshwater microalgae Scenedesmus quadricauda was studied using a set of biophysical and mathematical methods. Chlorophyll a fluorescence transients were recorded both in cell suspensions and at the level of single cells after incubation at copper concentrations of 0.1–10 $\mu$M under light and dark conditions. It was found that copper has a dose-dependent effect on the photosynthetic apparatus of microalgae. At low copper concentration (0.1 $\mu$M), a stimulating effect on a number of studied parameters was observed, whereas significant disruption of Photosystem II activity was detected at 10 $\mu$M. The method of analyzing fluorescence of single cells proved to be more sensitive compared to traditional suspension measurements, allowing the detection of heterogeneous cellular responses to the toxicant. Analysis of chlorophyll a fast fluorescence kinetics showed that the JIP-test parameters $\delta_{Ro}$ and $\varphi_{Ro}$ were the most sensitive to copper exposure and were significantly different from the control when exposed not only to high but also to medium (1 $\mu$M) copper concentrations. The decrease in photochemical activity of cells during light incubation was less pronounced compared to dark conditions. The application of data normalization to optical density at $\lambda = 455$ nm significantly increased the sensitivity of the method and accuracy of result interpretation. The use of L1-regularization (LASSO) by the least angles method (LARS) for the spectral multi-exponential approximation of the fluorescence transients allowed us to reveal their temporal characteristics. Mathematical analysis of the obtained data suggested that copper exposure leads to increased non-photochemical quenching of fluorescence, which serves as a protective mechanism for dissipating excess excitation energy. The revealed heterogeneity of cellular responses to copper action may have important ecological significance, ensuring the survival of part of the population under stress conditions. The obtained data confirm the promise of using fluorescent analysis methods for early diagnosis of heavy metal stress effects on photosynthesizing organisms.

  6. Epifanov A.V., Tsybulin V.G.
    Regarding the dynamics of cosymmetric predator – prey systems
    Computer Research and Modeling, 2017, v. 9, no. 5, pp. 799-813

    To study nonlinear effects of biological species interactions numerical-analytical approach is being developed. The approach is based on the cosymmetry theory accounting for the phenomenon of the emergence of a continuous family of solutions to differential equations where each solution can be obtained from the appropriate initial state. In problems of mathematical ecology the onset of cosymmetry is usually connected with a number of relationships between the parameters of the system. When the relationships collapse families vanish, we get a finite number of isolated solutions instead of a continuum of solutions and transient process can be long-term, dynamics taking place in a neighborhood of a family that has vanished due to cosymmetry collapse.

    We consider a model for spatiotemporal competition of predators or prey with an account for directed migration, Holling type II functional response and nonlinear prey growth function permitting Alley effect. We found out the conditions on system parameters under which there is linear with respect to population densities cosymmetry. It is demonstated that cosymmetry exists for any resource function in case of heterogeneous habitat. Numerical experiment in MATLAB is applied to compute steady states and oscillatory regimes in case of spatial heterogeneity.

    The dynamics of three population interactions (two predators and a prey, two prey and a predator) are considered. The onset of families of stationary distributions and limit cycle branching out of equlibria of a family that lose stability are investigated in case of homogeneous habitat. The study of the system for two prey and a predator gave a wonderful result of species coexistence. We have found out parameter regions where three families of stable solutions can be realized: coexistence of two prey in absence of a predator, stationary and oscillatory distributions of three coexisting species. Cosymmetry collapse is analyzed and long-term transient dynamics leading to solutions with the exclusion of one of prey or extinction of a predator is established in the numerical experiment.

    Views (last year): 12. Citations: 3 (RSCI).
  7. Zhdanova O.L., Kolbina E.A., Frisman E.Y.
    Evolutionary effects of non-selective sustainable harvesting in a genetically heterogeneous population
    Computer Research and Modeling, 2025, v. 17, no. 4, pp. 717-735

    The problem of harvest optimization remains a central challenge in mathematical biology. The concept of Maximum Sustainable Yield (MSY), widely used in optimal exploitation theory, proposes maintaining target populations at levels ensuring maximum reproduction, theoretically balancing economic benefits with resource conservation. While MSYbased management promotes population stability and system resilience, it faces significant limitations due to complex intrapopulation structures and nonlinear dynamics in exploited species. Of particular concern are the evolutionary consequences of harvesting, as artificial selection may drive changes divergent from natural selection pressures. Empirical evidence confirms that selective harvesting alters behavioral traits, reduces offspring quality, and modifies population gene pools. In contrast, the genetic impacts of non-selective harvesting remain poorly understood and require further investigation.

    This study examines how non-selective harvesting with constant removal rates affects evolution in genetically heterogeneous populations. We model genetic diversity controlled by a single diallelic locus, where different genotypes dominate at high/low densities: r-strategists (high fecundity) versus K-strategists (resource-limited resilience). The classical ecological and genetic model with discrete time is considered. The model assumes that the fitness of each genotype linearly depends on the population size. By including the harvesting withdrawal coefficient, the model allows for linking the problem of optimizing harvest with the that of predicting genotype selection.

    Analytical results demonstrate that under MSY harvesting the equilibrium genetic composition remains unchanged while population size halves. The type of genetic equilibrium may shift, as optimal harvest rates differ between equilibria. Natural K-strategist dominance may reverse toward r-strategists, whose high reproduction compensates for harvest losses. Critical harvesting thresholds triggering strategy shifts were identified.

    These findings explain why exploited populations show slow recovery after harvesting cessation: exploitation reinforces adaptations beneficial under removal pressure but maladaptive in natural conditions. For instance, captive arctic foxes select for high-productivity genotypes, whereas wild populations favor lower-fecundity/higher-survival phenotypes. This underscores the necessity of incorporating genetic dynamics into sustainable harvesting management strategies, as MSY policies may inadvertently alter evolutionary trajectories through density-dependent selection processes. Recovery periods must account for genetic adaptation timescales in management frameworks.

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